"It is now run and predominantly owned by chief executive Jay Graber as a US public benefit corporation."

It's that predominantly that's the tell. The Guardian has no more idea who owns which and how many shares as anyone else, because it's a privately held corporation. Yeah, i'm pretty sure, and when they go public, we'll all be positive of it. If Dorsey wasn't the largest shareholder, they would have been feeding THAT out to the press, instead of the incessant every present PR line "stepped down from the board" and "closed his account" lol

jack dorsey is the largest shareholder in bluesky because he founded it and has never transferred or given up his shares in it