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The national guard is part of the military, so funded and supported by the feds. Unlike normal army units though, each state or territory has its own national guard unit under the command of the governor. The intention is to give each state the power to quickly respond to emergency situations without needing federal approval. They're the successors to the old state militias, but have much stronger federal ties now.
What's the chain of command?
Does it stop at the Governor, or does it go from the Governor to the President?
Do they swear an allegiance to the state Constitution, or to the Federal Constitution?
They swear allegiance to the federal constitution, the governor, and the president. With the president being CIC and having power over the governor and the constitution having power overall. So in theory, the governor cannot give you orders that defy the president, and not even the president can give force you to comply with an unlawful order. It would take some serious stones to defy orders though